Dogs were the first of any species that people domesticated, and they have been a constant part of human life for millennia. Domesticated species are the plants and animals that have evolved to live alongside humans, providing nearly all of our food and numerous other benefits. Dogs provide protection, hunting assistance, companionship, transportation and even wool for weaving blankets.

Dogs evolved from gray wolves, but scientists debate exactly where, when and how many times dogs were domesticated. Ancient DNA evidence suggests that domestication happened twice, in eastern and western Eurasia, before the groups eventually mixed. That blended population was the ancestor of all dogs living today.

Molecular clock analysis of the DNA from hundreds of modern and ancient dogs suggests they were domesticated between around 20,000 and 22,000 years ago, when large ice sheets covered much of Eurasia and North America. The first dog identified in the archaeological record is a 14,000-year-old pup found in Bonn-Oberkassel, Germany, but it can be difficult to tell based on bones whether an animal was an early domestic dog or a wild wolf.

Despite the shared history of dogs and wolves, scientists have long thought these two species rarely mated and gave birth to hybrid offspring. As an evolutionary biologist and a molecular anthropologist who study domestic plants and animals, we wanted to take a new look at whether dog-wolf hybridization has really been all that uncommon.

Dogs are not exactly descended from modern wolves. Rather, dogs and wolves living today both derive from a shared ancient wolf population that lived alongside woolly mammoths and cave bears.

In most domesticated species, there are often........

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